1996 Diploma in Biochemistry, Eberhard Karls University Tübingen, Faculty of Chemistry, D. 2000 PhD thesis in Biochemistry, Ludwig-Maximilians-University Munich, Faculty of Chemistry, D. 2000 Postdoc, Max Planck Institute for Biochemistry (Martinsried, D). Till 2004 Postdoc, GSF National Research Center, (Neuherberg, D), Institute of Human Genetics. Till 2006 Postdoc, GSF National Research Center, (Neuherberg, D), Institute of Toxicology. 2006 Postdoc, Institute Gustave Roussy (Paris, F). Since 2007 to present: Research Group leader "Mitochondrial Toxicology", Institute of Molecular Toxicology and Pharmacology, Helmholtz Center Munich (Neuherberg, D). 2011 Habilitation in Toxicology, Technical University Munich, Medical faculty (D). Since 2017 to present: Research Group leader, Institute of Toxicology and Environmental Hygiene, Technical University of Munich, Germany. 2018 Professor in Toxicology, Technical University Munich, Medical faculty (D).